Treasure Boxes (Japanese: たからばこ Treasure Box) are items in the Pokémon Mystery Dungeon series, that first appeared in Explorers of Time and Explorers of Darkness. They can be dropped from defeated Pokémon or found on the floor.
In Explorers of Time, Darkness, and Explorers of Sky, Treasure Boxes start appearing once the player goes on the Fogbound Lake expedition, and can be opened at Xatu Appraisal located in Treasure Town for 150 
. At the end of certain dungeons, player is guaranteed to find one to six Deluxe Boxes that can contain rare items such as stat boosters or high-rarity exclusive items depending on the dungeon. The boxes dropped by enemies have random contents: usually a ☆ exclusive item (if the enemy's evolutionary family has one, it is likely to be chosen), sometimes instead a reward for a normal mission for that specific dungeon floor. The specific odds of a Treasure Box and whether it will contain an exclusive item or not is species-specific. The enemies will never drop a Treasure Box during a Special Episode or in Star Cave.
In Explorers of Sky, Secret Stairs found in postgame dungeons can lead to Secret Rooms with Treasure Boxes; these however have dungeon-specific item pools.
In the Pokémon Mystery Dungeon games for WiiWare, Treasure Boxes are unlocked after reaching second rank and can appear across all versions linked to the save file in any dungeon.
In Pokémon Mystery Dungeon: Gates to Infinity, the boxes can be opened by Rampardos Box Buster for 150 
. Unlike in previous games, Gummis do not exist and Max Elixirs are often obtained instead, although it is still possible to find other items such as held items, and Gold Bars in amounts from one to five. Treasure Boxes can also contain Team Skills. The Team Skill, Collector, enables the player to find more Treasure Boxes from defeated enemies. There is a 15 floor DLC Map named Skill Treasury that enables more boxes to be dropped by defeated Pokémon as well. Boxes can also be found in chambers that require Keys and/or a specific Pokémon type to enter. These chambers can be found with Upgrade 2 of the Detour Guide. Upgrade 3 of the Detour Guide also sells the keys required to enter these chambers for 1650 . These keys can also be bought by exchanging 5 Gold Bars at Glorious Gold in Post Town. Treasure Boxes are stored in a space separate from items in the Deposit Box.
In Pokémon Super Mystery Dungeon, the boxes can be opened at Rockin' Lock in Lively Town. Blue-colored boxes are the most common and can be opened for 150 
while the rarer red-colored boxes can be opened for 300 . Like the previous game, Treasure Boxes are stored in their own space in the Deposit Box.
In Pokémon Mystery Dungeon: Rescue Team DX, all Treasure Boxes in the player's Toolbox are automatically opened upon exiting a dungeon. Treasure Boxes cannot be deposited in Kangaskhan Storage.
In all games, contents of the boxes are predetermined at the moment it's generated, meaning that opening the box will always have the same result.
. At the end of certain dungeons, player is guaranteed to find one to six Deluxe Boxes that can contain rare items such as stat boosters or high-rarity exclusive items depending on the dungeon. The boxes dropped by enemies have random contents: usually a ☆ exclusive item (if the enemy's evolutionary family has one, it is likely to be chosen), sometimes instead a reward for a normal mission for that specific dungeon floor. The specific odds of a Treasure Box and whether it will contain an exclusive item or not is species-specific. The enemies will never drop a Treasure Box during a Special Episode or in Star Cave.
. Unlike in previous games, Gummis do not exist and Max Elixirs are often obtained instead, although it is still possible to find other items such as held items, and Gold Bars in amounts from one to five. Treasure Boxes can also contain Team Skills. The Team Skill, Collector, enables the player to find more Treasure Boxes from defeated enemies. There is a 15 floor DLC Map named Skill Treasury that enables more boxes to be dropped by defeated Pokémon as well. Boxes can also be found in chambers that require Keys and/or a specific Pokémon type to enter. These chambers can be found with Upgrade 2 of the Detour Guide. Upgrade 3 of the Detour Guide also sells the keys required to enter these chambers for 1650 
while the rarer red-colored boxes can be opened for 300 
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