In the core series games
If held by a Sharpedo, allows it to Mega Evolve into Mega Sharpedo if its Trainer has a Key Stone. Each Trainer can only Mega Evolve one Pokémon per battle.
Unaffected by item negating effects (such as Embargo and Klutz). Item manipulating effects (such as Trick and Fling) cannot give Sharpedo a Sharpedonite or remove a Sharpedonite from Sharpedo.
Outside of battle
If held by a Sharpedo, allows it to Mega Evolve into Mega Sharpedo when it performs a Spectacular Talent during a Pokémon Contest Spectacular while its Trainer has a Key Stone. Sharpedo will remain a Mega Sharpedo until the Talent Round ends.
Pokémon holding Sharpedonite in Pokémon Omega Ruby and Alpha Sapphire cannot be traded to Pokémon X and Y.
| One of a variety of mysterious Mega Stones. Have Sharpedo hold it, and this stone will enable it to Mega Evolve during battle.
In spin-off games
|| This section is incomplete.|
Please feel free to edit this section to add missing information and complete it.
In Pokémon Shuffle, the Sharpedonite allows the player to Mega Evolve Sharpedo.
The player can obtain the Sharpedonite by defeating Mega Sharpedo in Stage 600 found on Prasino Woods.
The Lucarionite was also available as a prize during the Mega Sharpedo Competitive Stage. This Competitive Stage ran on the Nintendo 3DS version from September 7 to 14, 2015; January 25 to February 2, 2016 and November 8 to 15, 2016. It ran on the Pokémon Shuffle Mobile from January 25 to February 2, 2016; and November 8 to 15, 2016.
The Sharpedonite enables the following Mega Evolution:
|| Mega Sharpedo
|| Adds one more Mega Sharpedo above the match.
Actually converts the tiles directly above all other Mega Sharpedo into a Mega Sharpedo. This effect can replace any disruption in the tile above and counts as erasing the content of the tile for damage, but does not affect black-cloud disruptions. Does nothing if the above tile is empty.
In the manga
In the Pokémon Adventures manga
In Omega Alpha Adventure 11, Archie was revealed to have obtained a Sharpedonite. After his apparent death, Amber took possession of the Mega Stone.
In other languages
|| 巨牙鯊進化石 Geuihngàhsā Jeunfa Sehk
|| 巨牙鯊進化石 / 巨牙鲨进化石 Jùyáshā Jìnhuà Shí
|| 샤크니아나이트 Sharknianite