Tapu Lele reappeared in Rising from the Ruins!, where it battled Ash's Rockruff and won easily. When Olivia and Gladion's Lycanroc brought it to be healed by the guardian deity. When Tapu Lele seemed to be incapable of completely healing Rockruff, it attacked the two Lycanroc until Ash arrived and stepped in to protect them. It then used Draining Kiss to drain energy from Ash and the two Lycanroc, allowing it to heal Rockruff. Tapu Lele also stayed to watch as Rockruff subsequently evolved into a Dusk FormLycanroc in the light of the setting sun, smiling happily as it the floated away afterwards.
In Revealing the Stuff of Legend!, Tapu Lele and the other guardian deities revealed that they were the ones who took Nebby to Melemele Island, where Ash would eventually find and take care of it. At the Altar of the Sunne, the four guardian deities performed a ritual that allowed Nebby to evolve into Solgaleo. When Ash received a Solganium Z from Nebby, he found that it was unable to fit in his regular Z-Ring. Tapu Koko then took the Z-Ring, and through a dance, it and the other guardian deities upgraded the Z-Ring into a Z-Power Ring so Ash could properly use Solgaleo's Z-Move, Searing Sunraze Smash. The four then dispersed after seeing Nebby carry Ash and his friends through the Ultra Wormhole.
Personality and characteristics
Tapu Lele seems to be a kind and caring Pokémon, as it healed Ash's wounds despite not knowing him. However, it also doesn't seem to mind battling much weaker opponents and hurting them in the process, as shown when it challenged and easily defeated Ash's Rockruff, acting gleefully happy the whole time.